Contents • • • • • • • • • Plot [ ] Down-on-his-luck Alpha Trax Joe Griffin () arrives at Waterloo Station in time to start his shift on one of the passenger train, which is scheduled to depart London at midnight under a full moon. A group of passengers is introduced, consisting of Kate (), Adrian (), Matthew (Amit Shah), Billy (Sam Gittens), Nina (), Paul () and Ged () and his wife Jenny ().
While he checks the passengers for tickets, he meets his unrequited love, the tea-trolley girl, Ellen (), who consoles him for being turned down for a promotion. An hour after departure, the train is travelling through a remote and forested part of the English countryside, in a. Here the train is forced to undergo an to prevent a derailment when a deer runs onto the track. The train driver Tony () goes outside to remove the body of the deer from the undercarriage. Whilst outside, Tony gets the deer's blood on him, and is suddenly eviscerated by a humanoid creature with phosphoric yellow eyes. Meanwhile, the passengers inside the stranded Alpha Trax are all initially strangers, from different walks of British life, are passively-aggressive towards each other, and demand that Joe get the train moving.
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The already strained situation is deepened when the train's fuel lines are severed and the radio antenna gets destroyed in a series of attacks, and subsequently the passengers are cut off from the rest of the world. After Joe discovers that Tony is missing, he tries to convince everyone to remain calm, but Adrian convinces everyone that the best choice is to disembark and walk towards the nearest station. Joe leads the group away from the train, but soon discovers the disemboweled body of Tony. They all immediately rush back into the train as the howls begin. Jenny is bitten by the werewolf as they try to drag her into the safety of the train. The group scramble to help Jenny, then sit around debating what it could have been that attacked and wounded Jenny.
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Kate goes to check on Nina when the werewolf attempts to get in through the door. Suddenly Nina's phone begins to ring, and just as she answers it the werewolf smashes through a window, grabbing her and dragging her through much to Kate's horror as Nina is heard screaming from the roof of the train and blood begins to pour down a window. Jenny's husband and old-timer passenger Ged () recalls a similar event happening in exactly the same spot - the Great Thornton Forest Rail Crash of 1963 - whereby all the passengers were found disemboweled and partially eaten the following morning. He motivates the remaining passengers to reinforce all the windows in the cabin to help prevent further break-ins by the werewolves.